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Department of Entomology, National Taiwan University

Core Faculty

Professor En-Cheng Yang
  • 發布單位:Department of Entomology

Research interests:

Neurophysiology, Insect vision



Lab: Insect Neurobiology Laboratory

Tel: +886-2-3366-9640



1990-1994 Ph D. National Australia University

1984-1988 MS. National Chung-Hsing University



2013/8-2016/7 Director, Mater Program of Plant Medicine, National Taiwan University

2013/8-2016/7 Chairman, Department of Entomology, National Taiwan University

2010/8-present Professor, Department of Entomology, National Taiwan University

2006/8-2010/7 Associate professor, National Taiwan University

2005-2006/7 Associate professor, National Chung-Hsing University

1999-2004 Assistant professor, National Chung-Hsing University

1996-1999 Postdoctoral Fellow, Institute of Molecular Biology, Academia Sinica

1994-1996 Postdoctoral Fellow, Centre for Visual Science, Australian National University

1990-1994 PhD student, Research School of Biological Sciences, Australian National University

1988-1989 Research Assistant, National Taiwan Normal University




  • Neurophysiology of insect color vision
  • Neurophysiology of insect form vision
  • Physiological function of nitric oxide in insect visual system
  • Stress physiology of honeybee

Application of pest control

  • Analysis of female attractant of Bactrocera dorsalis
  • Physical lure-and-kill equipments of Phyllotreta striolata in organic farming
  • Color preference as the method for pest control
  • Application of Wireless sensory network (WSN) in insect pest control and early warning system for quarantine purpose

Visual model

  • Computer image simulation in insect vision




Invertebrate Neurobiology

Ecology of Vision

Insect Physiology

Fundamental Entomology (part)

Insect Behavior (part)



Referred full-length articles


Papers published in academic journals

(*: corresponding author; SCI: journal in the list of scientific citation index)


1. Yang E.-C.* and Osorio D. (1991) Spectral sensitivities of photoreceptors and lamina monopolar cells in the dragonfly, Hemicordulia tau. J. Comp. Physiol. A 169: 663-670. [SCI]

2. Yang E.-C. and Osorio D.* (1996) Chromatic vision and photopic monopolar cells in the dragonfly lamina. J. Comp. Physiol. A 178: 543-550. [SCI]

3. Yang E.-C.* and Maddess T. (1997) Orientation-sensitive neurons in the brain of the honeybee (Apis mellifera). J. Insect Physiol. 43: 329-336. [SCI]

4. Maddess T.,* Davey M..P. and Yang E.-C. (1999) Discrimination of complex textures by bees. J. Comp. Physiol. A 184: 107-117. [SCI]

5. Tsai R.-S., Yang E.-C., Wu C.-Y., Tseng H.-K. and Chow Y.-S.* (1999) A potent sex attractant of the male tea tussock moth, Euproctis pseudoconspersa (Strand) (Lepidoptera; Lymantridae) in Taiwan: Field and EAG responses. Zool. Studies 38(3): 301-306. [SCI]

6. Stavenga D.G.,* Kinoshita M., Yang E.-C., Arikawa K. (2001) Retinal regionalization and heterogeneity of butterfly eyes. Naturwissenschaften 88: 477-481. [SCI]

7. Tso I.M.*, Tai P.I., Ku C.S. Kuo C.H. and Yang E.C.* (2002) Color-associated foraging success and population genetic structure in a polymorphic predator Nephila maculata (Araneae: Tetragnathidae). Animal Behaviour 63: 175-182. [SCI]

8. Hsieh Y.C., Yang E.C., Hsu E.L., Chow Y.S. and Kou R.* (2002) Voltage-dependent calcium channels in the corpora allata of adult male loreyi leafworm Mythimna loreyi. Insect Biochem. Molec. Biol. 32 (5): 547-557. [SCI]

9. Yang E.C.*, Lee D.W. and Wu W.Y. (2003) Action spectrum of phototactic responses of the flea beetle, Phyllotreta striolata. Physiol. Entomol. 28: 362-367. [SCI]

10. Tso I.M., Lin C.W. and Yang E.C.* (2004) Colourful orb-weaving spiders, Nephila pilipes, through a bee’s eyes. J. Exp. Biol. 207: 2631-2637. [SCI]

11. Yang E.C.*, Lin H.C. and Hung Y.S. (2004) Patterns of chromatic information processing in the lobula of the honeybee, Apis mellifera L. J. Insect Physiol. 50: 913-925. [SCI]

12. Tso I.M.*, Liao C.P., Huang R.P. and Yang E.C. (2006) Function of being pretty in web spiders: attracting prey or camouflaging oneself? Behavioral Ecology 17: 606-613. [SCI]

13. Yang E.C., Yang M.M.*, Liao L.H., Wu W.Y., Chen T.W. and Chen D.M. (2006) Non-destructive quarantine technique using soft X-ray imaging. Formosan Entomologist 26: 171-186.【榮獲96年度台灣昆蟲學會最佳論文獎】

14. Wu W.Y., Chen Y.B. and Yang E.C.* (2007) The chromatic cues to trap the oriental fruit fly, Bactrocera dorsalis. J.Insect Physiol. 53 :509-516. [SCI]

15. Chuang C.Y., Yang E.C. and Tso I.M*. (2007) Diurnal and nocturnal prey luring of a colorful predator. J. Exp. Biol. 210: 3830-3837. [SCI]

16. Chuang C.Y., Yang E.C. and  Tso I.M.* (2008) Deceptive color signaling in the night: a nocturnal predator attract prey with visual lures. Behavioral Ecology 19:237-244. [SCI]

17. Jiang J.A., Chang H.Y., Wu K.H., Ouyang C.S., Yang M.M., Yang E.C., and Lin T.T*. (2008) An adaptive image segmentation algorithm for X-ray quarantine insepection of selected fruits. Computers and Electronics in Agriculture 60: 190-200. [SCI]

18. Jiang J.A., Tseng C.L., Lu F., Yang E.C.*, Wu Z.S., Chen C.P., Lin S.H., Lin K.C., Liao C.S. (2008) A GSM-based remote wireless automatic monitoring system for field information: A case study for ecological monitoring of the oriental fruit fly, Bactrocera dorsalis (Hendel). Computers and Electronics in Agriculture 62: 243-259. [SCI]

19. Chen Y.L., Hung Y.S.and Yang E.C.* (2008) Biogenic amine level changes in the brains of stressed honeybees. Arch. Insect Biochem. and Physiol. 68: 241-250. [SCI]

20. Chio E.H. and Yang E.C.* (2008) A bioassay for natural insect repellents. J. of Asia-Pacific Entomol. 11: 225-227.

21. Yang E.C.*, Chuang Y.C., Chen Y.L. and Chang L.H. (2008) Abnormal foraging behavior induced by sublethal dosage of imidacloprid in the honey bee (Hymenoptera: Apidae). J. Econ. Entomol. 101: 1743-1748. [SCI]

22. Chen C.P., Chuang C.L., Tseng C.L., Yang E.C., Liu M.Y., Jiang J.A.* (2009) A Novel Energy-Efficient Adaptive Routing Protocol for Wireless Sensor Network. Journal of Chinese Society of Mechanical Engineers, 30: 59-65. [SCI]

23. Fan C.M., Yang E.C., Tso I.M.* (2009) Hunting efficiency and predation risk shapes the color-associated foraging traits of a predator. Behavioral Ecology 20: 808-816. [SCI]

24. Chiao C.C., Wu W.Y., Chen S.H., Yang E.C.* (2009) Visualization of the spatial and spectral signals of orb-weaving spiders, Nephila pilipes, through the eyes of a honeybee. Journal of Experimental Biology 212: 2269-2278 [SCI]

25. Jiang J.A.*, Chen C.P., Chuang C.L., Lin T.S., Tseng C.L., Yang E.C.,Wang Y.C. (2009) CoCMA: Energy-efficient coverage control in cluster-based wireless sensor networks using a memetic algorithm. Sensors 9: 4918-4940. [SCI]

26. Liu W.Y., Yang E.C.*, Lee S.J. (2009) The action spectrum of phototactic responses of Forcipomyia (Lasiohelea) taiwana (Shiraki) (Diptera: Ceratopogonidae). Formosan Entomologist 29: 61-71 (in Chinese)

27. Yao M.C., Lee C.Y., Yang E.C., Lu K.H.* (2009) Varieties and population fluctuations of stored-product insects in various rice storehouses monitored with light traps. Formosan Entomologist 29: 225-237. (in Chinese)

28. Liu W.Y., Lee S.J., Yang E.C.* (2009) Evaluation for attractiveness of four chemicals to the biting midge, Forcipomyia taiwana (Diptera: Ceratopogonidae). Journal of the American Mosquito Control Association 25(4): 448-455. [SCI]

29. Cheng R.C., Yang E.C., Lin C.-P. Herberstein M.E., Tso I.M.* (2010) Insect form vision as one potential shaping force of spider web decoration design. Journal of Experimental Biology  213: 759-768. [SCI]

30. Chuang C.L., Ouyang C.S., Lin T.T., Yang M.M., Yang E.C., Huang T.W., Kuei C.F., Luke A. Jiang J.A.* (2011) Automatic X-ray quarantine scanner and pest infestation detector for agricultural products. Computers and Electronics in Agriculture 77: 41-59. [SCI]

31. Okuyama T., Yang E.C., Chen C.P., Lin T.S., Chuang C.L., Jiang J.A.* (2011) Using automated monitoring systems to uncover pest population dynamics in agricultural fields. Agriculture System 104: 666-670. [SCI]

32. Wang T.S., Tung G.S.*, Yang E.C., Yang M.M. (2011) A preliminary study of controlling Quadrastichus erythrinae Kim on heritage coral trees with trunk injection. Formosan Entomology 31: 281-286.

33. Hsu P.S., Yang E.C.* (2012)  The critical cue in pattern discrimination for the honey bee: color or form? Journal of Insect Physiology 58: 934-940. [SCI]

34. Chen C., Yang E.C., Jiang J.A., Lin T.T*. (2012) An imaging system for monitoring the in-and-out activity of honey bees. Computers and Electronics in Agriculture 89:100-109. [SCI]

35. Yang E.C.*, Chang H.C., Wu W.Y., Chen Y.W. (2012) Impaired olfactory associative behavior of honeybee workers due to contamination of imidacloprid in the larval stage. PLoS ONE 7: e49472. [SCI]

36.  Jiang J.A., Lin T.S., Yang E.C., Tseng C.L., Chen C.P., Yen C.W., Zheng X.Y., Liu C.Y., Liu R.H., Chen Y.F., Chang W.Y., Chuang C.L.* (2012) Application of a web-based remote agro-ecological monitoring system for observing spatial distribution and dynamics of Bactrocera dorsalis in fruit orcahrds. Precision Agriculture [SCI]


37. Ho J.Z.*, Hu J.H., Yang E.C., Wu C.H., Yang P.S. (2012) The distribution, life cycle and flash behaviour of the firefly Pyrocoelia praetexta Olivier (Coleoptera: Lampyridae) in the Alishan area of Taiwan. Lampyrid 2:37-47.

38. Chen P.J., Arikawa K., Yang E.C.* (2013) Diversity of photoreceptors and spectral opponency in the compound eye of golden birdwing butterfly, Troides aeacus formosanus. PLoS ONE 8(4): e62240. doi:10.1371/journal.pone.0062240 [SCI]

39. Chio E.H., Yang E.C., Huang H.T., Hsu E.L., Chen C.R., Huang C.G., Huang R.N.* (2013) Toxicity and repellence of Taiwanese indigenous djulis, Chenopodium formosaneum, against Aedes albopictus (Diptera: Culicidae) and Forcipomyia taiwana (Diptera: Ceratopogonidae). J Pest Sci (DOI 10.1007/s10340-013-0500-3) (Chio and Yang are equal contributors) 86: 705-712. [SCI]

40. Tsai Z.M., Jau P.H., Kuo N.C., Kao J.C., Lin K.Y., Chang F.R., Yang E.C., Wang H.* (2013) A high range accuracy and high sensitivity harmonic radar using pulse pseudorandom code for bee searching. IEEE Trans. Microwave Theory and Tech. 61: 666-675. [SCI]

41. Jau P.H., Tsai Z.M., Kuo N.C., Kao J.C., Lin K.Y., Chang F.R., Yang E.C., Wang H.* (2014) Signal processing for harmonic pulse radar based on spread spectrum Technology. IET Radar, Sonar & Navigation. 8: 242-250. [SCI] (The IET Premium Awards 2015)

 42. Chen Y., Shih M.C., Wu M.H., Yang E.C., Chi K.J.* (2014) Underwater attachment using hairs: the functioning of spatula and sucker setae from male diving beetles. J R Soc Interface 20140273. [SCI]

 43. Tsai Y.L., Li C.W.*, Hong T.M., Ho J.Z., Yang E.C., Wu W.Y., Margaritondo G., Hsu S.T., Ong E.B.L., Hwu Y. (2014) Firefly ling flashing: Oxygen supply mechanism. Physical Rev Letters 113. 258103 [SCI]

44. Chiang W.Y., Wu M.H., Wu K.L., Lin M.H., Teng H.H., Tsai Y.F., Ko C.C., Yang E.C., Jiang J.A., Barnett L.R., Chu K.R.* (2014) A microwave applicator for uniform irradiation by circularly polarized waves in an anechoic chamber. Review of Scientific Instruments 85: 084703. [SCI]

45. Chuang C.L., Yang E.C., Tseng C.L., Chen C.P., Lien G.S., Jiang J.A.* (2014) Toward anticipating pest responses to fruit farms: Revealing factors influencing the population dynamics of the Oriental Fruit Fly via automatic field monitoring. Computers and Electronics in Agriculture 109: 148-161. [SCI]

46. Ou Yang T.H., Yang E.C., Jiang J.A., Lin T.T.* (2015) Mosquito vector monitoring system based on optical wingbeat classification. Computers and Electronics in Agriculture 118: 47-55. [SCI]

47. Peng Y.C. and Yang E.C.* (2016) Sublethal dosage of imidacloprid reduces the microglomerular density of honey bee mushroom bodies. Scientific Reports 6, 19298; doi: 10.1038/srep19298. [SCI]

48. Chen P.J., Awata H., Matsushita A., Yang E.C., Arikawa K*. (2016) Extreme spectral richness in the eye of the Common Bluebottle butterfly, Graphium sarpedon. Frontiers in Ecology and Evolution 4:18; doi:10.3389/fevo.2016.00018. [SCI]

49. Jiang J.A.*, Wang C.H., Chen C.H., Liao M.S., Su Y.L., Chen W.S., Huang C.P., Yang E.C., Chuang C.L. (2016) A WSN-based automatic monitoring system for the foraging behavior of honey bees and environmental factors of beehives. Computers and Electronics in Agriculture 123: 304–318. [SCI]

50. Liang C.H., Chuang C. L., Jiang J.A., Yang E.C.* (2016) Magnetic Sensing through the abdomen of the honey bee. Scientific Reports 6, 23657 (doi: 10.1038/srep23657) [SCI]

51. Chen Y.W*., Wu P.S., Yang E.C., Nai Y.S., Haung Z.Y. (2016) The impact of pyriproxyfen on the development of honey bee (Apis mellifera L.) colony in field. Joural of Asia-Pacific Entomology 19:589-594. (doi:10.1016/j.aspen.2016.06.005)[SCI]

52. Hsu M.L., Liu T.H., Yang T.C., Jhan H.C., Wang H., Chang F.R., Lin K.Y., Yang E.C., Tsai Z.M.* (2016) Bee searching radar with high transmit-receive isolation using pulse pseudorandom code. IEEE Transactions on Microwave Theory and Techniques 64:4324-4335. [SCI]

53. Hu Y.T., Wu T.C., Yang E.C., Wu P.C., Lin P.T. Wu Y.L.* (2017) Regulation of genes related to immune signaling and detoxification in Apis mellifera by an inhibitor of histone deacetylation. Scientific Reports 7, 41255 doi:10.1038/srep41255 [SCI]

54. Wu M.C., Chang Y.W., Lu K.H., Yang E.C.* (2017) Gene expression changes in honey bees induced by sublethal imidacloprid exposure during the larval stage. Insect Biochemistry and Molecular Biology 88:12-20. [SCI]

55. Peng Y.C., Yang E.C.* (2017) Reply to ‘Pitfalls of using confocal-microscipy based automated quantification of synaptic complexes in honeybee mushroom bodies (response to Peng and Yang 2016)’. Scientific Reports 7:11286. [SCI]

56. Pisa L., Goulson D., Yang E.C., Gibbons D., Sánchez-Bayo F., Mitchell E., Aebi A., van der Sluijs J., MacQuarrie C., Giorio  C., Long E.Y., McField E., Bijleveld van Lexmond M., Bonmatin J.M.* (2017)  An update of the Worldwide Integrated Assessment (WIA) on systemic insecticides. Part 2: Impacts on organisms and ecosystems. Environmental Science and Pollution Research DOI: 10.1007/s11356-017-0341-3 [SCI]

57. Owens A.C.S., Meyer-Rochow V.B., Yang E.C.(2018) Short- and mid-wavelength artificial light influences the flash signals of Aquatica ficta flash signals (Coleoptera: Lampyridae). PLoS ONE 13(2): e0191576 ( [SCI]

58. Pao S.H., Tsai P.Y., Peng C.I., Chen P.J., Tsai C.C., Yang E.C., Shih M.C., Chen J., Yang J.Y., Chesson P., and Sheue C.R. *. 2018. Lamelloplasts and Minichloroplasts in Begoniaceae: Iridescence and Photosynthetic Functioning. Journal of Plant Research 131: 655-670 (doi: 10.1007/s10265-018-1020-2). (cover page article) [SCI]

59. Hu Y.T., Tang C.K., Wu P.C., Yang E.C., Tsai C.C., Wu Y.L.* (2018) Histone deacetylase inhibitor-treatment restores memory-related gene expression and learning ability in neonicotinoid-treated Apis mellifera. Insect Molecular Biology 27: 512-521. (doi: 10.1111/imb.12390) [SCI]

60. Chen Y.R., Tzeng D., Owens A.C.S, Wu C.H., Hsiao C.Y., Tang H.C., Zhong S., Yang E.C.* (2018) The impact of artificial light on firefly larvae transcriptome. Formosan Entomology 38: 63-72. (in Chinese)

61.  Ngo T.N., Wu K.C., Yang E.C., Lin T.D* (2019) A Real-Time Imaging System for Multiple Honey Bee Tracking and Activity Monitoring. Computers and Electronics in Agriculture [SCI]

62. Tai K.C., Shrestha M.,  Dyer A.G., Yang E.C., Wang C.N.* (2020) Floral colour diversity: how are signals shaped by elevational gradient on the tropical-subtropical mountainous island of Taiwan? Frontiers in Plant Science[SCI]

63. Hsieh H.W., Chen S.C., Yang E.C., Hang S., Hsu C.C. Kou R.*(2021) Social interactions upregulate hemolymph tryptophan and tyrosine levels in the male lobster cockroach. Hormones and Behavior 130 104935 [SCI]

64. Chen Y.R., Wei W.L., Tzeng D. T.W., Owens A.C.S., Tang H.C., Wu C.S., Lin S.S., Zhong S., Yang E.C.* (2021) The effect of artificial light at night (ALAN) on gene expression of Aquatica ficta firefly larvae. Environmental Pollution 281:116944 [SCI]

65. Chen Y.R., Tzeng D.T.W. Ting C. Hsu P.S., Wu T.H., Zhong S.*, Yang E.C.*(2021) Missing nurse bees - Early transcriptomic switch from nurse bee to forager induced by sublethal imidacloprid. Frontiers in Genetics [SCI]

66. Ngo T.N., Rustia D.J.A, Yang E.C., Lin T.T. *(2021) Honey bee colony population daily loss rate forecasting and early warning method using temporal convolutional networks. Sensors 21: 3900 [SCI]

67. Ngo T.N., Wu K.C., Yang E.C., Lin T.D.* (2021) Automated Monitoring and Analyses of Honey Bee Pollen Foraging Behavior Using a Deep Learning-Based Imaging System. Computers and Electronics in Agriculture 187: 106239  [SCI]

68. Chen C.W., Whiting M.J.*, Yang E.C., Lin S.M.* (2021) Do I stay or do I go? Shifts in perch use by lizards during twilight suggests anticipatory behaviour. Biology Letters 17: 20210388. [SCI]

69. Chen Y.R., Tzeng D.T.W., Yang E.C.* (2021) Chronic effects of imidacloprid on honey bee worker development—molecular pathway perspectives. International Journal of Molecular Sciences 22: 11835 [SCI]


Articles presented in conference or published in non-academic journals

1. 楊恩誠、洪于善(2001)「色誘」昆蟲的理論基礎與應用。跨世紀台灣昆蟲學研究之進展研討會專刊 pp.69-77。

2. 楊恩誠、黃上銓、陳怡伶、桂佳鳳(2002)蜜蜂之趨光行為-對比感度與視神經機制。2002蜜蜂生物學研討會專刊 pp.67-78。

3. 楊恩誠、李德威 (2003) 色光在害蟲防治上的應用。農業世界雜誌 243:22-25。

4. 楊曼妙、楊正澤、楊恩誠 (2004) 植物隱匿性害蟲入侵可能途徑與檢測之簡介。植物重要防疫檢疫害蟲診斷鑑定研習會 (四) pp. 107-117。

5. 楊恩誠、詹美鈴 (2005) 談昆蟲複眼中的形與色。科學月刊 36(4): 292-297。

6. 楊恩誠、詹美鈴 (2005) 花非花,霧非霧。2004科學週「形」特展專刊-形的密碼。國科會出版。

7. Lin T.T., Chang H.Y., Wu K.H., Jian J.A., Ouyang C.S. Yang M.M. & Yang E.C. (2005) An Adaptive Image Segmentation Algorithm for X-Ray Quarantine Inspection of Selected Fruits. (Paper presented in 2005 ASAE Annual International Meeting, 17 - 20 July 2005).

8. 楊曼妙、楊恩誠、林達德、江昭皚、陳子偉 (2005) X光自動化檢疫系統之研發與應用。動植物防疫檢疫季刊 3: 61-63。

9. Lin T.T., Jian J.A., Ouyang C.S., Chang H.Y., Yang M.M. & Yang E.C. (2005) Integration of An Automatic X-ray Scanning System for Fruit Quarantine. (Paper presented in The Eight International Conference on Automation Technology Conference) Proceedings of Automation 2005: 1-6.

10. Ouyang C.S., Jiang J.A., Wu C.H., Yang P.S., Yang E.C. & Lin T.T. (2006) Development of the automatic image processing system for behavior of firefly luminescence. (Paper presented in the 3rd international Symposium on Machinery and Mechatronics Agricultural and Biosystems Engineering (ISMAB)) 23-25 November 2006, Seoul Korea. Proceeding of 3rd ISMAB 2006:  443-449.

11. Jiang J.A., Ouyang C.S., Lin T.T., Chang H.Y., Yang M.M., Yang E.C., Kuei J.F. & Chen T.W. (2006) Application of LabView platform to X-ray automatic quarantine system for fruits. (Paper presented in the 3rd international Symposium on Machinery and Mechatronics Agricultural and Biosystems Engineering (ISMAB)) 23-25 November 2006, Seoul Korea. Proceeding of 3rd ISMAB 2006: 747-754.

12. Chuang C.L., Ouyang C.S., Lin T.T., Yang M.M., Yang E.C., Chang H.Y., Kuei J.F. Chen T.W. & Jian J.A. (2006) An improved morphological-based approach for X-ray quarantine inspection of selected fruits. (Paper presented in the 3rd international Symposium on Machinery and Mechatronics Agricultural and Biosystems Engineering (ISMAB)) 23-25 November 2006, Seoul Korea. Proceeding of 3rd ISMAB 2006:  755-761.

13. 莊育禎、楊恩誠 (2007)低劑量益達胺對蜜蜂採蜜行為之影響。第五屆海峽兩岸蜜蜂生物學研討會。

14. 黃澤偉、楊曼妙、楊恩誠、江昭皚、林達德、陳子偉 (2007) X光自動化檢疫系統之研發與應用。植物重要防疫檢疫害蟲診斷鑑定研習會 (七) pp. 59-68。

15.  楊恩誠、江昭皚、曾傳蘆、盧福明、陳子偉、陳家榜、劉錕、林詩翔、林辰膺、曾主平、林子翔、廖誌聖、司仕豪、嚴崇瑋 (2008) 無線感測器網路技術在植物防檢疫之應用。動植物防疫檢疫季刊 17: 74-75。

16.  楊恩誠 (2008) 蜜蜂怎麼不見了? 麥克Do科學 25: 3。

17. 江昭皚、盧福明、楊恩誠、曾傳蘆 (2008) 無線感測器網路技術在農業害蟲監測之應用。農政與農情 194: 73-76。

18. 江昭皚、盧福明、楊恩誠、曾傳蘆、廖國基、嚴崇瑋 (2009)東方果實蠅生態監測與預警系統。自動化科技學會會刊 2009. 04: 54-65。

19. 楊恩誠 (2009) 蜜蜂與牠的視界。自然通訊 22: 29-32。

20. 楊恩誠、陳宏源 (2010) 昆蟲機師的完美著陸。科學發展月刊 449: 36-41。

21. 楊恩誠、王庭碩、陳宏源 (2010) 驅蚊及誘蚊相關防治技術。環境用藥管理及病媒防治技術研討會。2010年6月3日。台北,台灣。

22.吳佩珊、張惠君、莊育禎、陳裕文、楊恩誠 (2010) 非致死劑量農藥對蜜蜂的影響。海峽兩岸第八屆蜜蜂與蜂產品研討會論文集 1-12。2010年8月12-13日。天水,甘肅,中國。

23. Yang E. C., Wu P. S., Chang H. C., Chen Y. W. (2010) Effect of sub-lethal dosages of insecticides on honeybee behavior and physiology. International Seminar on enhancement of Functional Biodiversity Relevant to Sustainable Food Production in ASPAC. November 8-12, 2010. Tsukuba International Congress Center, Tsukuba, Ibaraki, Japan.

24. 楊恩誠 (2011) 益達胺對蜜蜂行為的影響。苗栗區農業專訊 55: 16-18。

25. 楊恩誠、江昭皚 (2012) 蜂之羅盤。科學人 122: 36-40。

26. Ho J.Z., Hu J.H., Yang E.C., Wu C.H., Yang P.S. (2012) The distribution, life cycle and flash behavior of the firefly Pyrocoelia praetexta Olivier (Coleoptera: Lampyridae) in the Alishan area of Taiwan. Lampyrid 2:37-47.

27. 楊恩誠、王庭碩 (2012) 台灣原生刺桐的隱形殺手—刺桐釉小蜂危害情形與緊急防治方法。臺大農業推廣通訊雙月刊  95: 1-5。

28. Jau P.H., Kuo N.C., Kao J.C., Lin K.Y., Chang F.R., Yang E.C., Wang H. (2012) A high range resolution 9.4/18.8 GHz harmonic radar for bees searching. Microwave Symposium Digest 1-3.

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